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Epilobiums
are often difficult to identify because many of them are interfertile,
i.e., they hybridize readily, or as Intermountain Flora states, there is "rampant hybridization in... Epilobium".
Therefore, Intermountain
Flora indicates, in about half of the species in our area,
"specific distinctions are... vague". The two species
shown below are in this vague group and are highly variable in many of
their growth characteristics.
Linnaeus
named this genus in 1753; the word "Epilobium" is from the
Greek for "upon a capsule" and refers to the long tubular
inferior ovary, shown extending to the left of the pinched area in the
left center of this photograph of Epilobium ciliatum.
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Epilobium
ciliatum (Willowherb) Onagraceae (Evening Primrose Family) Foothills, montane.
Roadsides, meadows. Summer. Epilobium ciliatum can grow in both moist and dry conditions, be two-to-six feet tall or just eight inches tall, and it is equally variable in leaf shape, amount of branching of the stem, and other characteristics. Even the silky fluff that carries the seeds can vary from white to dingy. The picture at left shows just the lower few inches of a plant over two feet tall. Leaves are very minutely toothed and stems often reddish. Constantine Rafinesque named this species in 1808. The species has had several dozen other names since then. "Ciliatum" is Latin for "hairy" and botanically refers to fine, marginal hairs -- which can be seen as a silvery glow along the edges of some of the seed pods and buds in the two pictures below.
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Epilobium
ciliatum (Willowherb) Onagraceae (Evening Primrose Family) Foothills, montane.
Roadsides, meadows. Summer. Pink flowers are most common for Epilobium ciliatum but white is almost as common and both pink and white flowers can appear on the same plant.
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Epilobium
ciliatum (Willowherb) Onagraceae (Evening Primrose Family) Foothills, montane.
Roadsides, meadows. Summer. The long, vertical seed pods shown in the above left picture will soon ripen and split in graceful arches exposing the soft tuft of hairs that are attached to and carry aloft tiny, brown, minutely ridged seeds. |
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Epilobium
hornemannii (Willowherb) Onagraceae (Evening Primrose Family) Montane, subalpine. Meadows, streamsides. Summer. Epilobium hornemannii flowers are tiny, pink/white/light-blue, and inconspicuous, but the plant is so common that it eventually attracts attention. The pictured specimen is lush; other Epilobium hornemannii (see below) may have far fewer and narrower leaves, fewer stems, and be shorter. Epilobium hornemannii is a highly variable plant. Exact identification is even more problematic since, as observed above, it also hybridizes . For instance, E. Hornemannii hybridizes readily with E. lactiflorum which occupies similar habitat as E. hornemannii and, according to William Weber, can be distinguished from E. hornemannii by its "several broad withered leaves [at the base of the stem] at flowering time." E. hornemannii has "small and inconspicuous or no withered leaves" at flowering time. Look for Epilobium hornemannii along moist montane and sub-alpine trails, near ponds, and along irrigation ditches. It is common for Willowherbs to have red stems and leaves in the spring and to return to these colors in the fall. Heinrich Reichenbach (1793-1879) named this species in 1824. Jens Hornemann was a Danish botanist of the early 1800s. (More biographical information.) |
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Epilobium hornemannii (Willowherb) Onagraceae (Evening Primrose Family) Montane, subalpine.
Meadows, streamsides. Summer. |
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Epilobium hornemannii (Willowherb) Onagraceae (Evening Primrose Family) Montane, subalpine.
Meadows, streamsides. Summer. |
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Epilobium hornemannii (Willowherb) Onagraceae (Evening Primrose Family) Montane, subalpine.
Meadows, streamsides. Summer. Epilobium hornemannii seed capsules (the dark vertical tubes in the lower and upper right) split in graceful arches (center) and the brown seeds imbedded in tiny white fluff are ready for a breeze. |
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